Tagus Basin
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The Tagus Basin is the
drainage basin A drainage basin is an area of land where all flowing surface water converges to a single point, such as a river mouth, or flows into another body of water, such as a lake or ocean. A basin is separated from adjacent basins by a perimeter, t ...
of the
Tagus River The Tagus ( ; es, Tajo ; pt, Tejo ; see below) is the longest river in the Iberian Peninsula. The river rises in the Montes Universales near Teruel, in mid-eastern Spain, flows , generally west with two main south-westward sections, to e ...
, which flows through the west of the
Iberian Peninsula The Iberian Peninsula (), ** * Aragonese and Occitan: ''Peninsula Iberica'' ** ** * french: Péninsule Ibérique * mwl, Península Eibérica * eu, Iberiar penintsula also known as Iberia, is a peninsula in southwestern Europe, defi ...
and empties into
Lisbon Lisbon (; pt, Lisboa ) is the capital and largest city of Portugal, with an estimated population of 544,851 within its administrative limits in an area of 100.05 km2. Grande Lisboa, Lisbon's urban area extends beyond the city's administr ...
. It covers an area of 78,467 km2, which is distributed 66% (55,645 km2) on
Spanish Spanish might refer to: * Items from or related to Spain: **Spaniards are a nation and ethnic group indigenous to Spain **Spanish language, spoken in Spain and many Latin American countries **Spanish cuisine Other places * Spanish, Ontario, Cana ...
territory and 34% on
Portuguese Portuguese may refer to: * anything of, from, or related to the country and nation of Portugal ** Portuguese cuisine, traditional foods ** Portuguese language, a Romance language *** Portuguese dialects, variants of the Portuguese language ** Portu ...
land (22,822 km2). It is the third largest basin in the Iberian Peninsula, after the
Douro Basin The Douro (, , ; es, Duero ; la, Durius) is the highest-flow river of the Iberian Peninsula. It rises near Duruelo de la Sierra in Soria Province, central Spain, meanders south briefly then flows generally west through the north-west part of ...
, with 98,258 km2, and the
Ebro Basin The Ebro Basin was a foreland basin that formed to the south of the Pyrenees during the Paleogene. It was also limited to the southeast by the Catalan Coastal Ranges. It began as a fully marine basin with connections to both the Atlantic Ocean and t ...
, with 82,587 km2.


Physical environment

The
Tagus The Tagus ( ; es, Tajo ; pt, Tejo ; see #Name, below) is the longest river in the Iberian Peninsula. The river rises in the Montes Universales near Teruel, in mid-eastern Spain, flows , generally west with two main south-westward sections ...
basin is one of the most important in the Peninsula, due to its extension and its flow, being the one with the highest population density in
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...
and the
peninsula A peninsula (; ) is a landform that extends from a mainland and is surrounded by water on most, but not all of its borders. A peninsula is also sometimes defined as a piece of land bordered by water on three of its sides. Peninsulas exist on all ...
. It is formed by an elongated surface with an east–west orientation, the
Tagus River The Tagus ( ; es, Tajo ; pt, Tejo ; see below) is the longest river in the Iberian Peninsula. The river rises in the Montes Universales near Teruel, in mid-eastern Spain, flows , generally west with two main south-westward sections, to e ...
flows from the Sierra de Albarracín, where it has its source, to the
estuary An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environment ...
, Mar de la Paja, next to
Lisbon Lisbon (; pt, Lisboa ) is the capital and largest city of Portugal, with an estimated population of 544,851 within its administrative limits in an area of 100.05 km2. Grande Lisboa, Lisbon's urban area extends beyond the city's administr ...
, through the center of the
Hesperian Massif The Iberian Massif (also Hesperian, Hesperic, Hercynian Massif or Meseta block) is the pre-Mesozoic core of the Iberian Peninsula. Roughly covering the western half of the Peninsula, it includes all its Precambrian and Paleozoic materials and it i ...
with a length of 910 km, in the Spanish area, 1092 km in total length. The basin is wedged between the
Central System The Central System, Spanish and pt, Sistema Central, is one of the main systems of mountain ranges in the Iberian Peninsula. The 2,592 m high Pico Almanzor is its highest summit. The Central System is located just north of the 40th parallel an ...
, to the north, the
Montes de Toledo The Montes de Toledo are one of the main systems of mountain ranges in the Iberian Peninsula. They divide the drainage basin of the Tagus from the basin of the Guadiana. The highest peak is high La Villuerca. Description The Montes de Toledo are ...
and
Sierra de Montánchez Sierra de Montánchez is a mountain range in the greater Montes de Toledo range, Spain. It is named after nearby Montánchez town, the capital of the Tierra de Montánchez comarca, located on its slopes. This range gives its name to the Mancomun ...
, to the south and the
Iberian System Iberian refers to Iberia. Most commonly Iberian refers to: *Someone or something originating in the Iberian Peninsula, namely from Spain, Portugal, Gibraltar and Andorra. The term ''Iberian'' is also used to refer to anything pertaining to the fo ...
, ( Serranía de Cuenca and Sierra de Albarracín), to the east; bordering to the north with the
Ebro , name_etymology = , image = Zaragoza shel.JPG , image_size = , image_caption = The Ebro River in Zaragoza , map = SpainEbroBasin.png , map_size = , map_caption = The Ebro ...
and
Douro The Douro (, , ; es, Duero ; la, Durius) is the highest-flow river of the Iberian Peninsula. It rises near Duruelo de la Sierra in Soria Province, central Spain, meanders south briefly then flows generally west through the north-west part of ...
basins; to the south with the Guadiana basin and to the east with the Ebro and
Júcar left The Júcar () or Xúquer () is a river on the Iberian Peninsula of Spain. The river runs for approximately 509 km from its source at Ojuelos de Valdeminguete, on the eastern flank of the Montes Universales, Sistema Ibérico. Its most i ...
basins. The western limit, as far as the
Spanish Spanish might refer to: * Items from or related to Spain: **Spaniards are a nation and ethnic group indigenous to Spain **Spanish language, spoken in Spain and many Latin American countries **Spanish cuisine Other places * Spanish, Ontario, Cana ...
area is concerned, is delimited by the Erjas and Sever rivers, which form the border with
Portugal Portugal, officially the Portuguese Republic ( pt, República Portuguesa, links=yes ), is a country whose mainland is located on the Iberian Peninsula of Southwestern Europe, and whose territory also includes the Atlantic archipelagos of ...
. In the interior of the area defined by these mountains and by the minor reliefs of the Hercynian massif, which completes the closure to the west, it is structured in a
graben In geology, a graben () is a depressed block of the crust of a planet or moon, bordered by parallel normal faults. Etymology ''Graben'' is a loan word from German, meaning 'ditch' or 'trench'. The word was first used in the geologic contex ...
filled by
Cenozoic The Cenozoic ( ; ) is Earth's current geological era, representing the last 66million years of Earth's history. It is characterised by the dominance of mammals, birds and flowering plants, a cooling and drying climate, and the current configura ...
materials, sands, clays, marls, gypsum and some limestone in the upper levels, which constitute horizons of silting up of the ancient lake that occupied the original depression. The mountain ridges of the Tagus basin only reach high altitudes in the
Central System The Central System, Spanish and pt, Sistema Central, is one of the main systems of mountain ranges in the Iberian Peninsula. The 2,592 m high Pico Almanzor is its highest summit. The Central System is located just north of the 40th parallel an ...
, especially in the middle and eastern sectors ( Sierra de Béjar,
Sierra de Gredos The Sierra de Gredos is a mountain range in central Spain that spans the provinces of Ávila, Salamanca, Cáceres, Madrid, and Toledo. It is part of the much larger Sistema Central of mountain ranges. Its highest point is Pico Almanzor, at 2,5 ...
and
Sierra de Guadarrama The Sierra de Guadarrama (Guadarrama Mountains) is a mountain range forming the main eastern section of the Sistema Central, the system of mountain ranges along the centre of the Iberian Peninsula. It is located between the systems Sierra de G ...
), where they frequently exceed 2000
meters above sea level Height above mean sea level is a measure of the vertical distance (height, elevation or altitude) of a location in reference to a historic mean sea level taken as a vertical datum. In geodesy, it is formalized as ''orthometric heights''. The comb ...
; in the
Iberian System Iberian refers to Iberia. Most commonly Iberian refers to: *Someone or something originating in the Iberian Peninsula, namely from Spain, Portugal, Gibraltar and Andorra. The term ''Iberian'' is also used to refer to anything pertaining to the fo ...
, only some peaks of the
Montes Universales Montes Universales is a long mountain range in the southeastern end of the Iberian System. Its highest point is the 1,935 m high summit known as Caimodorro. The 1,830 m high Muela de San Juan is another important peak. Administratively, the Mon ...
exceed 1800
m.a.s.l. Height above mean sea level is a measure of the vertical distance (height, elevation or altitude) of a location in reference to a historic mean sea level taken as a vertical datum. In geodesy, it is formalized as ''orthometric heights''. The com ...
, while in the
Montes de Toledo The Montes de Toledo are one of the main systems of mountain ranges in the Iberian Peninsula. They divide the drainage basin of the Tagus from the basin of the Guadiana. The highest peak is high La Villuerca. Description The Montes de Toledo are ...
the altitudes are considerably lower. The
altitude Altitude or height (also sometimes known as depth) is a distance measurement, usually in the vertical or "up" direction, between a reference datum and a point or object. The exact definition and reference datum varies according to the context ...
of the interior depression is much lower, although very variable, decreasing rapidly from the extreme northeast to the western edge, thus, while in the plains of
La Alcarria The Alcarria ( es, La Alcarria) is a natural region in Castile (Spain). It is located mainly in Guadalajara Province but also overlaps those of Cuenca and Madrid. Its principal attractions are its fauna and flora and it is noted for its honey a ...
the heights are close to 1000 m.a.s.l, in
Aranjuez Aranjuez () is a city and municipality of Spain, part of the Community of Madrid. Located in the southern end of the region, the main urban nucleus lies on the left bank of Tagus, a bit upstream the discharge of the Jarama. , the municipality h ...
they drop below 500 m.a.s.l, in
Navalmoral de la Mata Navalmoral de la Mata is a Municipalities in Spain, municipality of Spain located in the province of Cáceres, autonomous community of Extremadura. Attached to the traditional ''comarca'' of Campo Arañuelo, the municipality lies on central-wester ...
to 300 and in the lands to the south of Coria to little more than 200 m.a.s.l.. Therefore, some of the tributaries of the middle sector of the
Tagus The Tagus ( ; es, Tajo ; pt, Tejo ; see #Name, below) is the longest river in the Iberian Peninsula. The river rises in the Montes Universales near Teruel, in mid-eastern Spain, flows , generally west with two main south-westward sections ...
have captured part of the original Douro basin by
headward erosion Headward erosion is erosion at the origin of a stream channel, which causes the origin to move back away from the direction of the stream flow, lengthening the stream channel.Essentials of Geology, 3rd Ed, Stephen Marshak It can also refer to ...
, favored by the greater
gradient In vector calculus, the gradient of a scalar-valued differentiable function of several variables is the vector field (or vector-valued function) \nabla f whose value at a point p is the "direction and rate of fastest increase". If the gradi ...
s determined by the altimetric difference between the two basins, the most characteristic examples being the
Alberche The Alberche is a river in the provinces of Ávila, Madrid and Toledo, central Spain. It begins its course at 1,800 m in Fuente Alberche, San Martín de la Vega del Alberche municipal term, Ávila Province. It forms the natural division between ...
and the Alagón. The network of tributary rivers of the Tagus is very dissymmetrical, those on the right bank are the ones that provide the most abundant flows, as they collect the contributions from the
Central System The Central System, Spanish and pt, Sistema Central, is one of the main systems of mountain ranges in the Iberian Peninsula. The 2,592 m high Pico Almanzor is its highest summit. The Central System is located just north of the 40th parallel an ...
and the
Iberian System Iberian refers to Iberia. Most commonly Iberian refers to: *Someone or something originating in the Iberian Peninsula, namely from Spain, Portugal, Gibraltar and Andorra. The term ''Iberian'' is also used to refer to anything pertaining to the fo ...
; the left tributaries are generally shorter and have a low flow, especially those that originate in the
Montes de Toledo The Montes de Toledo are one of the main systems of mountain ranges in the Iberian Peninsula. They divide the drainage basin of the Tagus from the basin of the Guadiana. The highest peak is high La Villuerca. Description The Montes de Toledo are ...
. The longest rivers in the Tagus basin are:


Biotic framework

The biotic framework of the ''Tagus Basin'', due to its
geology Geology () is a branch of natural science concerned with Earth and other astronomical objects, the features or rocks of which it is composed, and the processes by which they change over time. Modern geology significantly overlaps all other Ear ...
,
geomorphology Geomorphology (from Ancient Greek: , ', "earth"; , ', "form"; and , ', "study") is the scientific study of the origin and evolution of topographic and bathymetric features created by physical, chemical or biological processes operating at or n ...
and
climatology Climatology (from Greek , ''klima'', "place, zone"; and , '' -logia'') or climate science is the scientific study of Earth's climate, typically defined as weather conditions averaged over a period of at least 30 years. This modern field of stud ...
, presents a great variety of
ecosystem An ecosystem (or ecological system) consists of all the organisms and the physical environment with which they interact. These biotic and abiotic components are linked together through nutrient cycles and energy flows. Energy enters the syste ...
s that include different
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
s and protected
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
; these ecosystems range from the high peaks of the mountain ranges of the
Central System The Central System, Spanish and pt, Sistema Central, is one of the main systems of mountain ranges in the Iberian Peninsula. The 2,592 m high Pico Almanzor is its highest summit. The Central System is located just north of the 40th parallel an ...
to the river valleys of the Upper Tagus or the alluvial plains of Toledo and Cáceres. The Tagus basin can be divided into two large biogeographical zones: the ''Luso-Extremaduran province'' (center-west), and the ''Castilian-Mestrazgo-Manchegan province'' (center-east), with its natural boundary at the confluence of the
Alberche The Alberche is a river in the provinces of Ávila, Madrid and Toledo, central Spain. It begins its course at 1,800 m in Fuente Alberche, San Martín de la Vega del Alberche municipal term, Ávila Province. It forms the natural division between ...
. In short, this is a climatic-lithological differentiation: the
limestone Limestone ( calcium carbonate ) is a type of carbonate sedimentary rock which is the main source of the material lime. It is composed mostly of the minerals calcite and aragonite, which are different crystal forms of . Limestone forms whe ...
sector in the center-east and the
siliceous Silicon dioxide, also known as silica, is an oxide of silicon with the chemical formula , most commonly found in nature as quartz and in various living organisms. In many parts of the world, silica is the major constituent of sand. Silica is one ...
soils in the center-west.


Vegetation

In general, the vegetation of the basin is as follows: in the high limestone mountain ranges, Serranía de Cuenca and Upper Tagus, there is a dense vegetation cover of needle-leaved forests dominated by ''
Pinus nigra ''Pinus nigra'', the Austrian pine or black pine, is a moderately variable species of pine, occurring across Southern Europe from the Iberian Peninsula to the eastern Mediterranean, on the Anatolian peninsula of Turkey, Corsica and Cyprus, as wel ...
'' and ''
Pinus sylvestris ''Pinus sylvestris'', the Scots pine (UK), Scotch pine (US) or Baltic pine, is a species of tree in the pine family Pinaceae that is native to Eurasia. It can readily be identified by its combination of fairly short, blue-green leaves and orang ...
'', pure or mixed; when climatic conditions are extreme, the pine forest is replaced by paramo vegetation or ''
Juniperus thurifera ''Juniperus thurifera'' (Spanish juniper) is a species of juniper native to the mountains of the western Mediterranean region, from southern France (including Corsica) across eastern and central Spain to Morocco and locally in northern Algeria. ...
'' juniper groves. Descending in
altitude Altitude or height (also sometimes known as depth) is a distance measurement, usually in the vertical or "up" direction, between a reference datum and a point or object. The exact definition and reference datum varies according to the context ...
and with the alternation of sunny and shady exposures, there are two types of subsclerophyllous forests: ''
Quercus faginea ''Quercus faginea'', the Portuguese oak, is a species of oak native to the western Mediterranean region in the Iberian Peninsula. Similar trees in the Atlas Mountains of northwest Africa are usually included in this species, or sometimes treate ...
'' Portuguese oak groves in the valley bottoms and shady slopes and occupying the so-called ''alcarrias'', (those of Torrecuadrada de los Valles,
Brihuega Brihuega is a municipality located in the Guadalajara (province), province of Guadalajara, Spain. According to the 2007 census (Instituto Nacional de Estadística (Spain), INE), the municipality had a population of 2,835 inhabitants. In 1710 a h ...
or
Jadraque Jadraque is a municipality of Spain located in the province of Guadalajara, Castilla–La Mancha. The municipality spans across a total area of 38.91 km2. As of 1 January 2020, it has a population of 1,370. History It is home to a medieval castl ...
are extensive), and mixed oak groves with Portuguese oaks and/or junipers in the sunny and rocky terrain, the main components of the
Tajuña The Tajuña (; from the Latin ''Tagonius'') is a river in central Spain, flowing through the provinces of Guadalajara and Madrid. It is a tributary of the river Jarama which in turn is a tributary of the Tagus. It rises in the Sierra de Solorio, n ...
valley. Occasionally, and on sandstone
lithology The lithology of a rock unit is a description of its physical characteristics visible at outcrop, in hand or core samples, or with low magnification microscopy. Physical characteristics include colour, texture, grain size, and composition. Lit ...
, there are oak or black pine forests, ''
Pinus pinaster ''Pinus pinaster'', the maritime pine or cluster pine, is a pine native to the south Atlantic Europe region and parts of the western Mediterranean. It is a hard, fast growing pine bearing small seeds with large wings. Description ''Pinus pinast ...
'', such as in the
Gallo Gallo may refer to: *Related to Gaul: ** Gallo-Roman culture **Gallo language, a regional language of France **Gallo-Romance, a branch of Romance languages **Gallo-Italic or Gallo-Italian language, a branch spoken in Northern Italy of the Romance ...
canyon,
Sigüenza Sigüenza () is a city in the Serranía de Guadalajara comarca, Province of Guadalajara, Castile-La Mancha, Spain. History The site of the ancient ''Segontia'' ('dominating over the valley') of the Celtiberian Arevaci, now called ('old town ...
or Poyatos, below these communities there are pure holm oak forests of continental character. The most frequent scrublands in this area are the
boxwoods ''Buxus'' is a genus of about seventy species in the family Buxaceae. Common names include box or boxwood. The boxes are native to western and southern Europe, southwest, southern and eastern Asia, Africa, Madagascar, northernmost South ...
, junipers, replacing the pine forests at the highest or most exposed heights and the ''erizals''. Underneath, there are steppe, ''
Cistus laurifolius ''Cistus laurifolius'', commonly called laurel-leaf cistus, laurel-leaved cistus or laurel-leaved rock rose, is a species of highly branched flowering evergreen shrub native plant, native to some areas around the Mediterranean Sea, Mediterranean. ...
'', sages and shrub pastures, as well as peat bogs and cervals of altitude or linked to hydromorphic
soil Soil, also commonly referred to as earth or dirt, is a mixture of organic matter, minerals, gases, liquids, and organisms that together support life. Some scientific definitions distinguish ''dirt'' from ''soil'' by restricting the former te ...
s. When the thermicity increases, either by the type of substrate or by the altitude, the pine forest becomes Aleppo pine, ''
Pinus halepensis ''Pinus halepensis'', commonly known as the Aleppo pine, also known as the Jerusalem pine, is a pine native to the Mediterranean region. Description ''Pinus halepensis'' is a small to medium-sized tree, tall, with a trunk diameter up to , exce ...
'', as in Sierra de Altomira, which is replaced by
kermes oak ''Quercus coccifera'', the kermes oak, is an oak bush in the ''Ilex'' section of the genus. It is native to the Mediterranean region and Northern African Maghreb, south to north from Morocco to France and west to east from Portugal to Cyprus an ...
, rosemary or even an
esparto grass Esparto, halfah grass, or esparto grass is a fiber produced from two species of perennial grasses of north Africa, Spain and Portugal. It is used for crafts, such as cords, basketry, and espadrilles. '' Stipa tenacissima'' and '' Lygeum spart ...
or esparto grove. To the north, the lithological change makes pine forests of Scots pine, ''
Pinus sylvestris ''Pinus sylvestris'', the Scots pine (UK), Scotch pine (US) or Baltic pine, is a species of tree in the pine family Pinaceae that is native to Eurasia. It can readily be identified by its combination of fairly short, blue-green leaves and orang ...
'', frequent, as in the Sierra de Pela and
Sierra de Ayllón The Sierra de Ayllón (Ayllón mountain chain) or Macizo de Ayllón (Ayllón massif) is a mountain chain of the Iberian Peninsula, belonging to the Sistema Central, of which it constitutes one of its easternmost spurs. It is located between the ...
, or the important pine forests of the Lozoya Valley and the upper basin of the
Guadarrama Guadarrama is a town and municipality in the Cuenca del Guadarrama comarca, in the Community of Madrid, Spain. Its population is 13,032 (winter, according to a 2006 census); the population swells to approximately 60,000 in summer. Its name co ...
and Aulencia rivers. Below, the domain of the ''
Quercus pyrenaica ''Quercus pyrenaica'', commonly known as Pyrenean oak, is a tree native to southwestern Europe and northwestern North Africa. Despite its common name, it is rarely found in the Pyrenees Mountains and is more abundant in northern Portugal and nort ...
'' Pyrenean oak groves begins, in
Sierra de Ayllón The Sierra de Ayllón (Ayllón mountain chain) or Macizo de Ayllón (Ayllón massif) is a mountain chain of the Iberian Peninsula, belonging to the Sistema Central, of which it constitutes one of its easternmost spurs. It is located between the ...
, Sierra de Somosierra,
Sierra de Guadarrama The Sierra de Guadarrama (Guadarrama Mountains) is a mountain range forming the main eastern section of the Sistema Central, the system of mountain ranges along the centre of the Iberian Peninsula. It is located between the systems Sierra de G ...
, part of
Sierra de Gredos The Sierra de Gredos is a mountain range in central Spain that spans the provinces of Ávila, Salamanca, Cáceres, Madrid, and Toledo. It is part of the much larger Sistema Central of mountain ranges. Its highest point is Pico Almanzor, at 2,5 ...
,
Sierra de Gata Sierra de Gata ( ext, Sierra e Gata) is one of the main mountain ranges in the Sistema Central, Spain. The highest point is Peña Canchera (1,592 m). Geography The Sierra de Gata is located in the northwest of the province of Cáceres, which is ...
and Sierra de Béjar; either by the action of man or by climatic-edaphic conditions, the oak grove is replaced by ''
Erica australis '' Erica australis'', the Spanish heath or Spanish tree heath, is a species of flowering plant in the family (biology), family Ericaceae, native to the western Iberian Peninsula (Portugal and Western Spain) and Northwest Africa (in Morocco). It i ...
'' heaths (Somosierra-Ayllón and center-west of Gredos, Béjar and Gata), or by ''
Cytisus scoparius ''Cytisus scoparius'' ( syn. ''Sarothamnus scoparius''), the common broom or Scotch broom, is a deciduous leguminous shrub native to western and central Europe. In Britain and Ireland, the standard name is broom; this name is also used for oth ...
'' or ''
Genista cinerea Genista is a genus of flowering plants in the legume family Fabaceae, native to open habitats such as moorland and pasture in Europe and western Asia. They include species commonly called broom, though the term may also refer to other genera, i ...
'' broom groves (Sierra de Guadarrama and eastern Gredos). At altitude, above the forest limit, the scrub is '' Cytisus oromediterraneus'' and '' Echinospartum barnadesii''. To the south, already in the middle of the evaporitic depression of the Tagus, the gypsophilous scrublands appear, so exclusive and rich in
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
and
vicariant Allopatric speciation () – also referred to as geographic speciation, vicariant speciation, or its earlier name the dumbbell model – is a mode of speciation that occurs when biological populations become geographically isolated from ...
species, which extend from the foothills of the Tagus in the Almoguera reservoir and reach past
Aranjuez Aranjuez () is a city and municipality of Spain, part of the Community of Madrid. Located in the southern end of the region, the main urban nucleus lies on the left bank of Tagus, a bit upstream the discharge of the Jarama. , the municipality h ...
and
Borox Borox is a municipality located in the Toledo (province), province of Toledo, Castile-La Mancha, Spain. According to the 2006 census (Instituto Nacional de Estadística (Spain), INE), the municipality had a population of 2640 inhabitants. Referen ...
, being also present in the lower basins of the
Jarama Jarama () is a river in central Spain. It flows north to south, and passes east of Madrid where the El Atazar Dam is built on a tributary, the Lozoya River. It flows into the river Tagus in Aranjuez. The Manzanares is a tributary of the Jarama ...
,
Tajuña The Tajuña (; from the Latin ''Tagonius'') is a river in central Spain, flowing through the provinces of Guadalajara and Madrid. It is a tributary of the river Jarama which in turn is a tributary of the Tagus. It rises in the Sierra de Solorio, n ...
or Manzanares. As a link between the gypsiferous substrates and the marly and calcareous ones, we find the esparto-rosemary and kermes oak groves. Descending in altitude from the
Central System The Central System, Spanish and pt, Sistema Central, is one of the main systems of mountain ranges in the Iberian Peninsula. The 2,592 m high Pico Almanzor is its highest summit. The Central System is located just north of the 40th parallel an ...
or the
Montes de Toledo The Montes de Toledo are one of the main systems of mountain ranges in the Iberian Peninsula. They divide the drainage basin of the Tagus from the basin of the Guadiana. The highest peak is high La Villuerca. Description The Montes de Toledo are ...
we enter the domain of the holm oak forest and the further west we move, the holm oak forest gives way to the
cork oak ''Quercus suber'', commonly called the cork oak, is a medium-sized, evergreen oak tree in the section ''Quercus'' sect. ''Cerris''. It is the primary source of cork for wine bottle stoppers and other uses, such as cork flooring and as the cores ...
grove, a typical
sclerophyll Sclerophyll is a type of vegetation that is adapted to long periods of dryness and heat. The plants feature hard leaf, leaves, short Internode (botany), internodes (the distance between leaves along the stem) and leaf orientation which is paral ...
ous community of clear Atlantic influence. These holm oak groves are replaced by various types of scrubland, from the gorse and thyme groves of the Alcarrias, passing through the
gum rockrose ''Cistus ladanifer'' is a species of flowering plant in the family Cistaceae. It is native to the western Mediterranean region. Common names include gum rockrose, labdanum, common gum cistus, and brown-eyed rockrose. Description It is a shrub ...
shrubs with Spanish lavender and
thyme Thyme () is the herb (dried aerial parts) of some members of the genus ''Thymus'' of aromatic perennial evergreen herbs in the mint family Lamiaceae. Thymes are relatives of the oregano genus ''Origanum'', with both plants being mostly indigenou ...
of the Sierra de Guadarrama or Montes de Toledo, by the broom groves of '' Cytisus multiflorus'', by the thickets of basil-leaved rock rose and other low rock-roses, or by the extensive retama groves of ''
Retama sphaerocarpa ''Retama'' (also known as ''rotem'', he, רותם) is a genus of flowering bushes in the legume family, Fabaceae. It belongs to the broom tribe, Genisteae. ''Retama'' broom bushes are found natively in North Africa, the Levant and some parts o ...
'' frequent in the plateau of Toledo and southern regions of the
Community of Madrid The Community of Madrid (; es, Comunidad de Madrid ) is one of the seventeen autonomous communities of Spain. It is located in the centre of the Iberian Peninsula, and of the Central Plateau (''Meseta Central''). Its capital and largest munici ...
. The
south South is one of the cardinal directions or Points of the compass, compass points. The direction is the opposite of north and is perpendicular to both east and west. Etymology The word ''south'' comes from Old English ''sūþ'', from earlier Pro ...
ern slopes of the Sierra de Guadarrama and Sierra de Gredos are home to important pine forests of cluster pine, ''
Pinus pinaster ''Pinus pinaster'', the maritime pine or cluster pine, is a pine native to the south Atlantic Europe region and parts of the western Mediterranean. It is a hard, fast growing pine bearing small seeds with large wings. Description ''Pinus pinast ...
'', stone pine, '' Pinus pinea'', and
relicts A relict is a surviving remnant of a natural phenomenon. Biology A relict (or relic) is an organism that at an earlier time was abundant in a large area but now occurs at only one or a few small areas. Geology and geomorphology In geology, a r ...
of black pine, ''
Pinus nigra ''Pinus nigra'', the Austrian pine or black pine, is a moderately variable species of pine, occurring across Southern Europe from the Iberian Peninsula to the eastern Mediterranean, on the Anatolian peninsula of Turkey, Corsica and Cyprus, as wel ...
'', the latter two species mainly in Gredos and in the Cofio-
Alberche The Alberche is a river in the provinces of Ávila, Madrid and Toledo, central Spain. It begins its course at 1,800 m in Fuente Alberche, San Martín de la Vega del Alberche municipal term, Ávila Province. It forms the natural division between ...
basin; among these stands alternate
Spanish chestnut ''Castanea sativa'', the sweet chestnut, Spanish chestnut or just chestnut, is a species of tree in the family Fagaceae, native to Southern Europe and Asia Minor, and widely cultivated throughout the temperate world. A substantial, long-lived ...
forests (Upper Hurdano,
Peña de Francia The Peña de Francia is a mountain that rises to , south of the Salamanca, Autonomous Region of Castile and León, Spain, whose peak belongs to the municipality of El Cabaco. Location It belongs to the municipality of El Cabaco, it is one of t ...
,
Las Batuecas Las Batuecas is a Spanish valley region of the Sierra de Francia in Salamanca Province, Castilla y León. It is located in the vicinity of La Alberca and is named after the river that runs through Las Batuecas. It has a monastery of cloistered ...
, Tiétar,
Valle del Jerte Valle del Jerte is a comarca and mancomunidad in the province of Cáceres, Spain. It contains the following municipalities: * Barrado * Cabezuela del Valle * Cabrero * Casas del Castañar * El Torno * Jerte * Navaconcejo Navaconcejo is ...
), '' Quercus broteroi'' Portuguese-oak groves (Upper Ibor, Almonte, Gébalo, Salor, etc.). To the west of Toledo and east of Cáceres, Campo Arañuelo, south of
Ávila Ávila (, , ) is a city of Spain located in the autonomous community of Castile and León. It is the capital and most populated municipality of the Province of Ávila. It lies on the right bank of the Adaja river. Located more than 1,130 m abov ...
, Tiétar valley, and almost all of
Extremadura Extremadura (; ext, Estremaúra; pt, Estremadura; Fala: ''Extremaúra'') is an autonomous community of Spain. Its capital city is Mérida, and its largest city is Badajoz. Located in the central-western part of the Iberian Peninsula, it ...
, there are holm oak and cork oak ''
dehesa A ''dehesa'' () is a multifunctional, agrosylvopastoral system (a type of agroforestry) and cultural landscape of southern and central Spain and southern Portugal; in Portugal, it is known as a ''montado''. Its name comes from the Latin 'defens ...
s'' or mixed masses of them. The ''dehesas'' with livestock use are a refuge of the floristic diversity, characteristic of the Iberian west. Another type of ''dehesas'', those of ash or oak (''
Fraxinus angustifolia ''Fraxinus angustifolia'', the narrow-leaved ash, is a species of ''Fraxinus'' native to central and southern Europe, northwest Africa, and southwest Asia.Flora Europaea''Fraxinus angustifolia''/ref>Rushforth, K. (1999). ''Trees of Britain and E ...
'' and ''
Quercus pyrenaica ''Quercus pyrenaica'', commonly known as Pyrenean oak, is a tree native to southwestern Europe and northwestern North Africa. Despite its common name, it is rarely found in the Pyrenees Mountains and is more abundant in northern Portugal and nort ...
'') are located in the foothills of the Sierra de Guadarrama and Sierra de Béjar.


Riverbank vegetation

In order to describe these plant communities of the Tagus basin, it is convenient to make a zoning that allows grouping areas that, while respecting the hydrological characteristics, present similar phytosociological characteristics.


Upper Tagus

The riparian vegetation of the Upper Tagus is characterized by the almost continuous presence of two types of willow groves of calcareous character, one of arboreal size that occupies the fresh banks with good soil in areas of medium and high mountains and another of shrubby character that usually occupies secondary channels, margins and alluvial beds with a high water table in areas of medium mountains and foothills. The first of the willow groves, generally corresponding to the ''Salicetum purpureo-albae'' association, is composed of one or two bands of vegetation, conditioned by the profile of the valley, where willows such as ''
Salix atrocinerea ''Salix atrocinerea'', commonly called grey willow or large gray willow, is a species of willow. It is a bush or small tree up to tall. As a pioneer species of willow, it quickly colonizes poor soils. Distribution The grey willow distribution ...
,
Salix alba ''Salix alba'', the white willow, is a species of willow native to Europe and western and central Asia.Meikle, R. D. (1984). ''Willows and Poplars of Great Britain and Ireland''. BSBI Handbook No. 4. .Rushforth, K. (1999). ''Trees of Britain an ...
, Salix fragilis'' or '' Salix neotricha'' dominate. They are usually accompanied by other tree species such as ''
Fraxinus angustifolia ''Fraxinus angustifolia'', the narrow-leaved ash, is a species of ''Fraxinus'' native to central and southern Europe, northwest Africa, and southwest Asia.Flora Europaea''Fraxinus angustifolia''/ref>Rushforth, K. (1999). ''Trees of Britain and E ...
,
Populus nigra ''Populus nigra'', the black poplar, is a species of cottonwood poplar, the type species of section ''Aigeiros'' of the genus ''Populus'', native to Europe, southwest and central Asia, and northwest Africa.Flora Europaea''Populus nigra''/ref> ...
'' and ''
alba ''Alba'' ( , ) is the Scottish Gaelic name for Scotland. It is also, in English language historiography, used to refer to the polity of Picts and Scottish people, Scots united in the ninth century as the Kingdom of Alba, until it developed i ...
,
Corylus avellana ''Corylus avellana'', the common hazel, is a species of flowering plant in the birch family Betulaceae. It is native to Europe and western Asia. It is an important component of the hedgerows that were the traditional field boundaries in lowland En ...
'' or ''
Cornus sanguinea ''Cornus sanguinea'', the common dogwood or bloody dogwood, is a species of dogwood native to most of Europe and western Asia, from England and central Scotland east to the Caspian Sea. It is widely grown as an ornamental plant. Description It ...
'', always surrounded by a thorny border of brambles and roses; within their courtship appear herbaceous plants such as ''
Equisetum ramosissimum ''Equisetum ramosissimum'' , known as branched horsetail, is a species of evergreen horsetail (genus ''Equisetum'', subgenus ''Hippochaete''). It is not the same species as ''Equisetum ramosissimum'' , which is a synonym of ''Equisetum giganteum' ...
,
Brachypodium sylvaticum ''Brachypodium sylvaticum'', commonly known as false-brome, slender false brome or wood false brome, is a perennial grass native to Europe, Asia and Africa. It has a broad native range stretching from North Africa to Eurasia. The bunchgrass is ...
'' or ''
Carex acutiformis ''Carex acutiformis'', the lesser pond-sedge, is a species of sedge. Description It grows up to tall, with leaves up to long and wide. Ecology It is native to parts of northern and western Europe, where it grows in moist spots in a number of ...
.'' More frequent and widely distributed in the Upper Tagus are the calcareous shrubby willow groves, ''alliance Salicion discolori-neotrichae''. They are generally dense formations, the typical wicker beds, which occupy the secondary channels, banks and alluvial beds of hard water rivers and more or less irregular regime, with floods and low water, withstanding floods well; its characteristic species are ''
Salix purpurea ''Salix purpurea'', the purple willow purpleosier willow or purple osier, is a species of willow native to most of Europe and western Asia north to the British Isles, Poland, and the Baltic States.Flora Europaea''Salix purpurea''/ref>Meikle, R. D ...
var. lambertiana'' and '' Salix elaeagnos subsp. angustifolia'', the latter dominating in the cooler areas. It is common to find meadows-rushes and even reed beds of the ''Molinio-Holoschoenion alliance'' and isolated specimens of ''
Populus nigra ''Populus nigra'', the black poplar, is a species of cottonwood poplar, the type species of section ''Aigeiros'' of the genus ''Populus'', native to Europe, southwest and central Asia, and northwest Africa.Flora Europaea''Populus nigra''/ref> ...
'' or ''
Populus alba ''Populus alba'', commonly called silver poplar,Webb, C. J.; Sykes, W. R.; Garnock-Jones, P. J. 1988: Flora of New Zealand. Vol. IV. Naturalised Pteridophytes, Gymnosperms, Dicotyledons. 4. Christchurch, New Zealand, Botany Division, D.S.I.R. si ...
'', as well as ''
Berberis vulgaris ''Berberis vulgaris'', also known as common barberry, European barberry or simply barberry, is a shrub in the genus ''Berberis'' native to the Old World. It produces edible but sharply acidic berries, which people in many countries eat as a ta ...
,
Viburnum lantana ''Viburnum lantana'', the wayfarer or wayfaring tree, is a species of '' Viburnum'', native to central, southern and western Europe (north to Yorkshire in England), northwest Africa, and southwestern Asia.Blamey, M. & Grey-Wilson, C. (1989). ' ...
'' or ''Ligustrum vulgare''. In other areas of the basin of lower altitude these willows form the first band of Gallery forest, riparian vegetation. Examples are some stretches of the Tagus in Morillejo, the Hoz Seca and Upper Cabrillas streams and numerous small riverbeds in this area.


The Tagus between Bolarque and Toledo

Downstream of the large reservoirs of Entrepeñas Reservoir, Entrepeñas and Buendía Reservoir, Buendía, the Tagus is a regulated river. In the first stretches, cultivated poplar groves dominate, but from the Bolarque Reservoir, Bolarque and Almoguera Reservoir, Almoguera reservoirs onwards, the riverbank vegetation changes drastically with respect to the upper stretch in several ways: * The main river and some of its tributaries have Hydraulic structure, hydraulic works that regulate their flow. * It crosses a wide area dominated by evaporitic substrates, gypsum, which provide certain conditions of xericity and salinity to the
soil Soil, also commonly referred to as earth or dirt, is a mixture of organic matter, minerals, gases, liquids, and organisms that together support life. Some scientific definitions distinguish ''dirt'' from ''soil'' by restricting the former te ...
. * The climate is more Mediterranean, with less annual rainfall and hotter and drier summers. * The riparian vegetation is reduced to a narrow band due to the agricultural use of the river meadows. Poplar groves, also called cottonwood groves, are usually dominated by ''
Populus alba ''Populus alba'', commonly called silver poplar,Webb, C. J.; Sykes, W. R.; Garnock-Jones, P. J. 1988: Flora of New Zealand. Vol. IV. Naturalised Pteridophytes, Gymnosperms, Dicotyledons. 4. Christchurch, New Zealand, Botany Division, D.S.I.R. si ...
,
Salix alba ''Salix alba'', the white willow, is a species of willow native to Europe and western and central Asia.Meikle, R. D. (1984). ''Willows and Poplars of Great Britain and Ireland''. BSBI Handbook No. 4. .Rushforth, K. (1999). ''Trees of Britain an ...
'' or ''
Populus nigra ''Populus nigra'', the black poplar, is a species of cottonwood poplar, the type species of section ''Aigeiros'' of the genus ''Populus'', native to Europe, southwest and central Asia, and northwest Africa.Flora Europaea''Populus nigra''/ref> ...
'', and are located on hydromorphic soils, rich in bases and deep, silty or silty-sandy in permanent Stream bed, streambeds or other forms of edaphic hydromorphism. When well preserved, it is an exuberant forest of both horizontal and vertical stratification, with well-developed arboreal, lianoid, shrub-spiny and herbaceous strata. The most frequent type of poplar grove in this area corresponds to the ''Rubio tinctorum-Populetum albae association'', where in addition to the aforementioned species there are other arboreal willows such as '' Salix fragilis, Ulmus minor'' or ''
Fraxinus angustifolia ''Fraxinus angustifolia'', the narrow-leaved ash, is a species of ''Fraxinus'' native to central and southern Europe, northwest Africa, and southwest Asia.Flora Europaea''Fraxinus angustifolia''/ref>Rushforth, K. (1999). ''Trees of Britain and E ...
''. In the more xerothermophilous areas, as in the Cedrón stream, headwaters of the Martín Román stream, Algodor River, Algodor river, Tagus river in Estremera, the poplar grove is accompanied by tamarisks, ''Tamarix gallica'' and ''Tamarix africana'', which become dominant in small meanders or abandoned riverbeds. The Martín Román stream supplies the Carrizal de Villamejor, one of the best preserved examples of helophytic vegetation, both in this Stream#Types, brook, as in other slopes to the Tagus, on the right (Borox and Guatén streams) the accumulation of salts causes the appearance of a halophilic tamarisk. In some streams, the riparian vegetation consists of tayaral with halophilic elements such as ''Caroxylon vermiculatum, Salsola vermiculata, Atriplex halimus, Sonchus maritimus'', etc. The edges with temporary waterlogging are where there is the greatest originality and floristic richness with communities of ''hygrohalophilic Chenopodioideae, quenopodiaceae'' such as ''Salicornia ramosissima, Suaeda splendens, Microcnemum coralloides'', these examples are located in the Cavina, Salinas and Cuevas streams between
Aranjuez Aranjuez () is a city and municipality of Spain, part of the Community of Madrid. Located in the southern end of the region, the main urban nucleus lies on the left bank of Tagus, a bit upstream the discharge of the Jarama. , the municipality h ...
and Yepes. Another increasingly scarce element are the elm trees, the best represented, although in a very precarious state of conservation, are found in the Algodor River, Algodor river, downstream of the Finisterre Reservoir, Finisterre reservoir, in the headwaters and slopes of the Martín Román stream where specimens of boxthorn, ''Flueggea tinctoria'' endemism luso-extremeño, are beginning to appear. Within the group of vines are frequent ''Humulus lupulus, Bryonia dioica, Dioscorea communis, Tamus communis, Rubia tinctorum'' and ''Vitis sylvestris''. The shrub stratum, when well preserved, is usually dense and rich in thorny plants such as ''Rubus, Rubus spp, Crataegus monogyna, Damson, Prunus insititia, Sambucus nigra'', etc.


Tajuña and Henares

These two rivers have similar characteristics: they originate in a calcareous environment and cross detrital soils in some part of their course. The main Tributary, tributaries are the: * Tajuña, Tajuña River: which receives water from the Ungría and Peñón rivers, and from the Matayeguas and San Andrés streams. * Henares, Henares River, which receives water from the Dulce, Salado, Hoz, Cercadillo, Frío, Cañamares, Bornova, Sorbe, Riatillo, Aliendre, Badiel, Torote rivers and Las Dueñas and Camarmilla streams. In all the rivers that originate in calcareous soils (Bornova, Cañamares, Tajuña, Henares, Cercadillo and Dulce) there is a calcareous shrubby scrubland, black scrubland, as the first band of vegetation of the ''Rubio coryliforii-Salicetum atrocinereae'' faciation. In this willow grove, which can reach arboreal size, there are numerous thorny plants, ''Prunus spinosa, Rubus spp., Rose ssp''. and scarce other shrubs, in some locations they are accompanied by Eurosiberian elements such as ''Populus tremula,
Corylus avellana ''Corylus avellana'', the common hazel, is a species of flowering plant in the birch family Betulaceae. It is native to Europe and western Asia. It is an important component of the hedgerows that were the traditional field boundaries in lowland En ...
, Rhamnus cathartica, Rhamnus catharticus'' and ''Ligustrum vulgare''. In some places they are even invaded by ''Quercus ilex, Ulmus minor'' and ''Genista scorpius''. Other more frequent species also appear in other areas of the basin such as '' Salix fragilis'' and ''
Salix purpurea ''Salix purpurea'', the purple willow purpleosier willow or purple osier, is a species of willow native to most of Europe and western Asia north to the British Isles, Poland, and the Baltic States.Flora Europaea''Salix purpurea''/ref>Meikle, R. D ...
'' var. ''lambertiana'', this is the case of the Upper Tajuña and some small adjacent watercourses (Prado stream). In the headwaters of the Sorbe, Lillas, Sonsaz, Riatillo and Frío rivers there are siliceous willow groves that can have arboreal habit in the Upper Sorbe or in the Riatillo. These are the same willow groves found further west in the Alberche, Jarama and Guadarrama basins, ''Rubio corylifolii-Salicetum atrocinereae'', dominated by ''
Salix atrocinerea ''Salix atrocinerea'', commonly called grey willow or large gray willow, is a species of willow. It is a bush or small tree up to tall. As a pioneer species of willow, it quickly colonizes poor soils. Distribution The grey willow distribution ...
'' interspersed with ''Betula pendula, Betula pubescens, Betula alba,
Pinus sylvestris ''Pinus sylvestris'', the Scots pine (UK), Scotch pine (US) or Baltic pine, is a species of tree in the pine family Pinaceae that is native to Eurasia. It can readily be identified by its combination of fairly short, blue-green leaves and orang ...
,
Quercus pyrenaica ''Quercus pyrenaica'', commonly known as Pyrenean oak, is a tree native to southwestern Europe and northwestern North Africa. Despite its common name, it is rarely found in the Pyrenees Mountains and is more abundant in northern Portugal and nort ...
, Taxus baccata, Ilex aquifolium, Fagus sylvatica'' or ''Populus tremula''. Under this canopy are usually found stands of ''Erica arborea'' and a notable representation of ferns, especially abundant, ''Urtica dioica''; it is also common to find stands of ''Fragaria vesca, Digitalis purpurea'' and ''Sorbus aucuparia''. When the direct influence of the altitude and the more humid climate of the headwaters of these rivers is abandoned, the sauceda is impoverished in tree species and if the conditions of xericity increase, specimens of ''Erica scoparia'' can be found, as in the Sorbe and the Riatillo. As important floristic curiosities, there are the oligotrophic alder groves of the Bornoba and Sorbe rivers upstream of the Alcorlo and Beleña reservoirs respectively, belonging to the association ''Galio broteriani-Alnetum glutinosae'', (the Gredense alder groves), and are composed by species such as ''Carex elata ssp. reuteriana, Erica arborea, Prunella vulgaris, Viola riviniana, Wahlenbergia hederacea, Prunus avium,'' etc. They are narrow woods along the river bed dominated by alder, ''Alnus glutinosa''. Sometimes there is a small band of shrubby willow grove in front, while towards the interior they are in contact with ash and Pyrenean oak groves. Below these alder groves, in the Henares and Sorbe rivers, there are mesotrophic alder groves of ''Salici rubentis-Alnetum glutosinae'', accompanied by ''
Salix alba ''Salix alba'', the white willow, is a species of willow native to Europe and western and central Asia.Meikle, R. D. (1984). ''Willows and Poplars of Great Britain and Ireland''. BSBI Handbook No. 4. .Rushforth, K. (1999). ''Trees of Britain an ...
'' and ''Salix rubens'' and often ''Tamarix gallica'', ''
Salix purpurea ''Salix purpurea'', the purple willow purpleosier willow or purple osier, is a species of willow native to most of Europe and western Asia north to the British Isles, Poland, and the Baltic States.Flora Europaea''Salix purpurea''/ref>Meikle, R. D ...
'' and other calcareous species such as ''Cephalanthera rubra,
Cornus sanguinea ''Cornus sanguinea'', the common dogwood or bloody dogwood, is a species of dogwood native to most of Europe and western Asia, from England and central Scotland east to the Caspian Sea. It is widely grown as an ornamental plant. Description It ...
'' or ''Humulus lupulus'', being replaced, when the original composition is altered, by willow and poplar groves with ''Salix salviifolia, Salix elaeagnos'' and ''
Salix purpurea ''Salix purpurea'', the purple willow purpleosier willow or purple osier, is a species of willow native to most of Europe and western Asia north to the British Isles, Poland, and the Baltic States.Flora Europaea''Salix purpurea''/ref>Meikle, R. D ...
.'' At lower altitudes, ash and poplar groves dominate, the former, with red osiers and dogwoods (''Cornus sanguineae-Fraxinetum angustifoliae''), are frequent in the lower reaches of the rivers Frío, Hoz, Dulce, Salado, Tajuña and the upper reaches of the Badiel, These ash woods prefer steep Topography, topographies that limit the rate of evapotranspiration and when well preserved it is a pluristrative forest with other tree species such as ''
Salix purpurea ''Salix purpurea'', the purple willow purpleosier willow or purple osier, is a species of willow native to most of Europe and western Asia north to the British Isles, Poland, and the Baltic States.Flora Europaea''Salix purpurea''/ref>Meikle, R. D ...
,
Cornus sanguinea ''Cornus sanguinea'', the common dogwood or bloody dogwood, is a species of dogwood native to most of Europe and western Asia, from England and central Scotland east to the Caspian Sea. It is widely grown as an ornamental plant. Description It ...
, Ligustrum vulgare, Prunus mahaleb, Sambucus nigra'', various brambles and hawthorns and other species typical of the surrounding communities such as ''
Quercus faginea ''Quercus faginea'', the Portuguese oak, is a species of oak native to the western Mediterranean region in the Iberian Peninsula. Similar trees in the Atlas Mountains of northwest Africa are usually included in this species, or sometimes treate ...
,
Juniperus thurifera ''Juniperus thurifera'' (Spanish juniper) is a species of juniper native to the mountains of the western Mediterranean region, from southern France (including Corsica) across eastern and central Spain to Morocco and locally in northern Algeria. ...
, Bupleurum rigidum,'' etc. It is worth mentioning the presence of tamarisk (''Tamaricetum gallicae'') which replaces the poplar and willow groves when thermoxericity is very pronounced or when there is a salinized Phreatic zone, water table. The second situation is described in some sections of the Salado, Hoz and Cercadillo rivers and the first is the most representative of the lower reaches of the Henares and Tajuña rivers. The best representation is the tayaral of Soto de Aldovea and the one that marks the lower Henares at its confluence with the Jarama, there are also remains of tayaral in the Tajuña. ''Tamarix gallica'' dominates, although there is also ''Tamarix africana'', and sporadically there are feet of ''
Salix alba ''Salix alba'', the white willow, is a species of willow native to Europe and western and central Asia.Meikle, R. D. (1984). ''Willows and Poplars of Great Britain and Ireland''. BSBI Handbook No. 4. .Rushforth, K. (1999). ''Trees of Britain an ...
, Ulmus minor'' and even ''
Populus alba ''Populus alba'', commonly called silver poplar,Webb, C. J.; Sykes, W. R.; Garnock-Jones, P. J. 1988: Flora of New Zealand. Vol. IV. Naturalised Pteridophytes, Gymnosperms, Dicotyledons. 4. Christchurch, New Zealand, Botany Division, D.S.I.R. si ...
''. If it is a typically xerophytic tayaral, it is accompanied by ''Artemisia campestris, Carlina corymbosa, Asteriscus aquaticus, Glycirrhiza glabra, Glycirhiza glabra,
Retama sphaerocarpa ''Retama'' (also known as ''rotem'', he, רותם) is a genus of flowering bushes in the legume family, Fabaceae. It belongs to the broom tribe, Genisteae. ''Retama'' broom bushes are found natively in North Africa, the Levant and some parts o ...
'' or ''Asparagus acutifolius.'' The remains of elm groves are also interesting, with a still important development in the Badiel, near the confluence with the Henares, in the middle reaches of the Camarmilla and Pantuerta streams. In a large part of its distribution area, the elm is replaced by a dense reedbed, as occurs in the Anchuelo stream or in the Camarmilla stream, or in the lower Tajuña.


Jarama and Guadarrama

These two rivers have similar characteristics in a large part of their course, although the lower section of the
Jarama Jarama () is a river in central Spain. It flows north to south, and passes east of Madrid where the El Atazar Dam is built on a tributary, the Lozoya River. It flows into the river Tagus in Aranjuez. The Manzanares is a tributary of the Jarama ...
crosses the Tagus evaporite trench. Both are born in a siliceous environment (
Sierra de Ayllón The Sierra de Ayllón (Ayllón mountain chain) or Macizo de Ayllón (Ayllón massif) is a mountain chain of the Iberian Peninsula, belonging to the Sistema Central, of which it constitutes one of its easternmost spurs. It is located between the ...
and
Sierra de Guadarrama The Sierra de Guadarrama (Guadarrama Mountains) is a mountain range forming the main eastern section of the Sistema Central, the system of mountain ranges along the centre of the Iberian Peninsula. It is located between the systems Sierra de G ...
), and flow through detritic soils of reeds (
Jarama Jarama () is a river in central Spain. It flows north to south, and passes east of Madrid where the El Atazar Dam is built on a tributary, the Lozoya River. It flows into the river Tagus in Aranjuez. The Manzanares is a tributary of the Jarama ...
) and sandy soils (Guadarrama) in some parts of their course. The main riverbeds of the Jarama basin (excluding the Henares and Tajuña rivers) are: Jaramilla, Lozoya (river), Lozoya, Valle, Puebla, Guadalix River, Guadalix, Veguillas, Navacerrada, Manzanares, Canencia, de Galga, de Vatorrón, de Viñuelas, Manina, Meaques, Culebro and de Trofas streams. And of the Guadarrama are: Aulencia river, Guatel river, Soto and Rielves streams. The upper Jarama and upper Guadarrama have the same arboreal and shrubby black willows as those described for the Sorbe and Riatillo, the best examples are found in the upper Jaramilla and Veguillas stream, with occasional presence of ''Betula alba, Taxus baccata,
Pinus sylvestris ''Pinus sylvestris'', the Scots pine (UK), Scotch pine (US) or Baltic pine, is a species of tree in the pine family Pinaceae that is native to Eurasia. It can readily be identified by its combination of fairly short, blue-green leaves and orang ...
, Sorbus aria'' and ''Ilex aquifolium'', the same occurs with the best preserved stands in the Valle river, Lozoya above the Pinilla reservoir, upper Guadalix or upper Manzanares (above the Santillana reservoir). In areas where the valley becomes very narrow, as in the Jaramilla, ''Erica arborea'' heath with ''
Cistus laurifolius ''Cistus laurifolius'', commonly called laurel-leaf cistus, laurel-leaved cistus or laurel-leaved rock rose, is a species of highly branched flowering evergreen shrub native plant, native to some areas around the Mediterranean Sea, Mediterranean. ...
'', replaces the willow grove. The arborescent ''Salix salviifolia'' willow groves of the upper Lozoya, downstream of the Pinilla dam and up to the Puentes Viejas reservoir, ''Salicito saviifolio-lambertianae subas. caricetosum'', in addition to the aforementioned willow, others such as ''
Salix atrocinerea ''Salix atrocinerea'', commonly called grey willow or large gray willow, is a species of willow. It is a bush or small tree up to tall. As a pioneer species of willow, it quickly colonizes poor soils. Distribution The grey willow distribution ...
, Salix triandra'' or '' Salix fragilis'' are observed, appearing occasionally alders, birches, Frangula alnus, alder buckthorns, Quercus pyrenaica, Pyrenean oaks or Quercus faginea, Portuguese oaks. As the riverbeds approach the outskirts of Madrid they are altered in all their composition, being more frequent the reeds and rushes that invade the riverbeds. Even so, there are still good examples of poplar groves with Salix alba, white willow and ash groves with willows, as in the poplar grove of the lower section of the Trofas Stream to the confluence with the Manzanares, the section of the Manzanares downstream of the Pardo reservoir or the final section at its confluence with the Jarama in Rivas-Vaciamadrid, where there are also abundant remains of elm groves. In these rivers, gallery ashes (''Salici saviifoliae-Fraxinetum angustifoliae'') are very frequent. This type of acidophilic ash grove is accompanied by a good number of ''Salix salviifolia'' as a consequence of the contact with the water of the riverbed and the shallowness of the water table.


Middle Tagus

This zone includes from the city of Toledo to the Valdecañas reservoir, here the Tagus is regulated practically continuously from the Villacañas reservoir to Alcántara Dam, Alcántara, near the border with
Portugal Portugal, officially the Portuguese Republic ( pt, República Portuguesa, links=yes ), is a country whose mainland is located on the Iberian Peninsula of Southwestern Europe, and whose territory also includes the Atlantic archipelagos of ...
. In this area there is a biogeographical change (entering the Portuguese-Extremeño domain) and lithological change, with metamorphic and plutonic substrates dominating, alternating with some sedimentary substrates, which give the
soil Soil, also commonly referred to as earth or dirt, is a mixture of organic matter, minerals, gases, liquids, and organisms that together support life. Some scientific definitions distinguish ''dirt'' from ''soil'' by restricting the former te ...
an acid or neutral character. The first half of this area stands out for the encasement of the riverbeds in the sedimentary basin of the Tagus, with fluvial terraces of easily eroded detrital materials. The riverside vegetation has been modified by the presence of poplar production. Alternating with these, one can still find poplar groves of ''
Populus alba ''Populus alba'', commonly called silver poplar,Webb, C. J.; Sykes, W. R.; Garnock-Jones, P. J. 1988: Flora of New Zealand. Vol. IV. Naturalised Pteridophytes, Gymnosperms, Dicotyledons. 4. Christchurch, New Zealand, Botany Division, D.S.I.R. si ...
'', including species such as ''
Fraxinus angustifolia ''Fraxinus angustifolia'', the narrow-leaved ash, is a species of ''Fraxinus'' native to central and southern Europe, northwest Africa, and southwest Asia.Flora Europaea''Fraxinus angustifolia''/ref>Rushforth, K. (1999). ''Trees of Britain and E ...
, Ulmus minor,'' etc. Also noteworthy are the island forests made up of willow groves and ash groves. In addition, the willows are usually accompanied by a band of ''Scirpus holoschoenus'' roundhead bulrush and another of ''Tamarix gallica'' French tamarix, in the Torcón stream we find the first important formations of tamujar, ''Flueggea tinctoria'', as riparian vegetation, increasing its presence the further west of the basin. At the headwaters of Chorro stream, a tributary of the Pusa, there is a sample of ''Betula pendula'' ssp. ''fontqueri'' birch (''Galium broterai-Betuletum parvibractaeatum''), accompanied by ''
Fraxinus angustifolia ''Fraxinus angustifolia'', the narrow-leaved ash, is a species of ''Fraxinus'' native to central and southern Europe, northwest Africa, and southwest Asia.Flora Europaea''Fraxinus angustifolia''/ref>Rushforth, K. (1999). ''Trees of Britain and E ...
, Sorbus torminalis, Ilex aquifolium'' and numerous scyphilous and nemoral plants (''Scilla ramburei, Galium broterianum, Poa nemoralis''). When this birch grove is altered, it is replaced by a willow grove with myrtle (''Frangulo alni-Myricaetum galeae''). In the headwaters of the Gévalo River, we can find remnants of ''Prunus lusitanica'' Portuguese laurel cherry shrubs, relict forests of the Cenozoic tropical flora, which can also be found in Extremadura in the upper, narrower and more sheltered stretches of the Ibor, Gualija and Viejas Rivers.


Tiétar and Alberche

These two rivers, of Gredense origin, give rise to some of the basins in which the riparian vegetation is best preserved, some sections being protected as River Reserves (Guadyervas) or as catalogued riverbanks (complete Tiétar,
Alberche The Alberche is a river in the provinces of Ávila, Madrid and Toledo, central Spain. It begins its course at 1,800 m in Fuente Alberche, San Martín de la Vega del Alberche municipal term, Ávila Province. It forms the natural division between ...
and tributaries of the right bank in Ávila). We are in the domain of the alder groves and shrubby mountain willow groves. The willow groves form narrow bands of ''
Salix atrocinerea ''Salix atrocinerea'', commonly called grey willow or large gray willow, is a species of willow. It is a bush or small tree up to tall. As a pioneer species of willow, it quickly colonizes poor soils. Distribution The grey willow distribution ...
'' with the presence of ''Betula alba, Taxus baccata, Ilex aquifolium,
Pinus nigra ''Pinus nigra'', the Austrian pine or black pine, is a moderately variable species of pine, occurring across Southern Europe from the Iberian Peninsula to the eastern Mediterranean, on the Anatolian peninsula of Turkey, Corsica and Cyprus, as wel ...
,
Pinus sylvestris ''Pinus sylvestris'', the Scots pine (UK), Scotch pine (US) or Baltic pine, is a species of tree in the pine family Pinaceae that is native to Eurasia. It can readily be identified by its combination of fairly short, blue-green leaves and orang ...
, Castanea sativa, Frangula alnus'', etc. An excellent example is the willow groves on the northern slope of the
Sierra de Gredos The Sierra de Gredos is a mountain range in central Spain that spans the provinces of Ávila, Salamanca, Cáceres, Madrid, and Toledo. It is part of the much larger Sistema Central of mountain ranges. Its highest point is Pico Almanzor, at 2,5 ...
that flow into the Alberche, such as Garganta Iruelas and Valsaína (they also include ''
Corylus avellana ''Corylus avellana'', the common hazel, is a species of flowering plant in the birch family Betulaceae. It is native to Europe and western Asia. It is an important component of the hedgerows that were the traditional field boundaries in lowland En ...
'' and ''Ulmus glabra''), Lanchamala and La Yedra. The willow grove of the Garganta stream (Serranillos-Navarrevisca) stands out with an excellent undergrowth of ''Viburnum opulus'' and ''Prunus padus''. In the higher areas, above the sauceda or where it cannot develop, ''Erica arborea'' heath can also be found as riparian vegetation. The alder groves can be divided into three types. On the one hand, the oligotrophic alder grove, very similar to the one that marks the upper Jarama, which is located in the Alberche river and its tributaries up to the Picadas reservoir. Behind this band, if the width of the valley allows it, an ash grove with willows ('' Salix fragilis'') can develop. This is the case of the Alberche, Cofio and Garganta Iruelas alder groves, the Alberche alder grove between Burgohondo and the tail of the Burguillo reservoir being one of the most important examples in the
Iberian Peninsula The Iberian Peninsula (), ** * Aragonese and Occitan: ''Peninsula Iberica'' ** ** * french: Péninsule Ibérique * mwl, Península Eibérica * eu, Iberiar penintsula also known as Iberia, is a peninsula in southwestern Europe, defi ...
. Finally, the ''Luso-Extremeña'' alder grove, where two differential species ''Osmunda regalis'' and ''Flueggea tinctoria'' are frequent, as well as the presence of ''Viburnum tinus, Erica lusitanica, Genista falcata'' and ''Acer monspessulanum''. These are the alder groves of the southern gorges of Gredos.


International Tagus

The end of the Tagus basin in Spain, constitutes the so-called ''international Tagus'', border for many kilometers between the two countries. Spanish tributaries in this area are the Aburrel, the Aurela and the Sever, as well as the Salor. In Portugal the Sever river and the Erges river. This area preserves very good examples of riparian forests, within the riverbed (in the channels that dry out in the Baseflow, low water) there is a tamujar sometimes accompanied by bog-myrtle, ''Myrica gale'', which gives way to an alder grove in permanent contact with the water, just behind this, and even alternating with it, there is a willow grove that further away from the bank connects with the poplar of ''
Populus nigra ''Populus nigra'', the black poplar, is a species of cottonwood poplar, the type species of section ''Aigeiros'' of the genus ''Populus'', native to Europe, southwest and central Asia, and northwest Africa.Flora Europaea''Populus nigra''/ref> ...
''. Where there is an alluvial plain there is also an ash grove (''Ficario ranunuculoidis-Fraxinetum angustifoliae''), behind it, a series of phreatophytic communities dominated by rushes of roundhead bulrush, ''Scirpus holoschoenus'', and a seasonal sub-humid grassland.


Fauna

The great diversity of relief and vegetation allows the existence of a rich and varied fauna. In the Tagus basin we can observe, within the group of vertebrates, about 66 species of mammals, 198 nesting birds, 26 reptiles, 18 amphibians and 29 fish, including numerous emblematic species of great value in the autonomous, state and international level. In addition to the fauna closely linked to aquatic ecosystems, the Tagus basin is home to the best populations of the black vulture, ''Aegypius monachus'', in Spain and Europe, as well as the Iberian imperial eagle, ''Aquila adalberti'', an Iberian endemism well represented in the basin, and finally the Iberian lynx, ''Lynx pardinus'', an emblematic species of the Spanish fauna and endemic to the Iberian Peninsula.


Macroinvertebrates

The so-called macroinvertebrates are represented by the taxa ''Insecta, Mollusca, Oligochaeta'' and ''Crustacean, Crustacea''. These species are adapted to a series of specific conditions of the aquatic environment, occupying different stretches of the river according to their habitat requirements and water purity. High mountain rivers and streams, areas of clean, oxygenated and cold waters, present a great variety and taxonomic richness with the usual presence of ''Plecoptera'' (families ''Perlidae'', ''Leuctridae'', etc.), ''Ephemeroptera'' (families ''Heptageniidae'', ''Ephemeridae'', ''Leptophlebiidae'', etc.) and ''Trichoptera'' (''Sericostomatidae'', ''Glossosomatidae'', ''Lepidostomatidae'', etc.) in their waters. The Mediterranean mountain has a lower diversity, although relatively high, being well represented some ''heteroptera'' and trichoptera adapted to low current and temperate waters (families ''Hydroptilidae, Hydroptylidae'', ''Rhyacophilidae'', etc.), the crustaceans ''Gammaridae'' typical in calcareous and mineralized areas and some plecoptera (family ''Nemouridae'') and ephemeroptera (families ''Caenidae'', ''Baetidae'', etc.). Macroinvertebrate taxa adapted to the aquatic environment present in the Tagus basin and included in the different red books or inventories that have some type of national or international protection: Within the Order (biology), order Odonata, Odonatos, three species included in the Habitats Directive, Habitat Directive are found in the ''Tagus basin'': Other invertebrates of interest in the ''Tagus basin'' are: Among the Mollusca, mollusks proposed by the Spanish Society of Malacology to be included in the National Catalogue of Threatened Species cited in the Tagus basin is included the ''hydroid, Neohoratia coronadoi'', located only in the
Jarama Jarama () is a river in central Spain. It flows north to south, and passes east of Madrid where the El Atazar Dam is built on a tributary, the Lozoya River. It flows into the river Tagus in Aranjuez. The Manzanares is a tributary of the Jarama ...
river, in the Canal de La Parra and in the Patones spring in the Community of Madrid. This Gastropoda, gastropod is proposed as ''endangered''.


Fish

In the ''Tagus basin'', 29 species of fish belonging to the families ''Anguillidae, Salmonidae, Esocidae, Cyprinidae, Cobitidae, Ictaluridae, Siluridae, Poeciliidae, Centrarchidae'' and ''Percidae'' have been cited. Of this group of species, 16 are native and 14 are introduced. Of the native species, 2 are endemic to Spain and 11 are endemic to the Iberian Peninsula. The headwaters of the main tributaries of the Tagus and the main course itself, the only fish species that inhabits is the brown trout, ''Salmo trutta'', a species that lives in fast, cold and well oxygenated waters. They are abundant in headwater areas, becoming scarcer as you descend in altitude. Downstream from the headwaters, conditions are allowing, together with the trout, the appearance of other species such as bermejuela, ''Chondrostoma arcasii'', ''colmilleja'', ''Cobitis paludica'' and ''Cobitis vettonica'', Northern Iberian Spined Loach, ''Cobitis calderoni'', Iberian nase, ''Chondrostoma polylepis'', ''madrilla'', ''Chondrostoma miegii'', Iberian gudgeon, ''Gobio lozanoi'', ''cacho'' ''Squalius pyrenaicus'', ''bordallo'' ''Squalius carolitertii'', ''calandino'' ''Squalius alburnoides''. The above species are all native to the peninsula or endemic to Spain (''Parachondrostoma arrigonis'' and ''Cobitis vettonica''). In slower and warmer waters, these species that become less abundant are joined by the ''Barbus bocagei'' and the ''Chondrostoma lemmingii'' and in lower areas by the Iberian barbel, ''Barbus comizo'' and the ''Barbus microcephalus''. Finally, the tench ''Tinca Tinca, Tinca tinca'', considered native to Spain due to the existence of fossil records from the Bronze Age, appears sporadically and occasionally in the middle and lower reaches of some rivers (Alagón, Jerte, Magasca, Guadiloba, Cuervo, etc.) and abundantly in ponds and dammed lagoons in the pasture oak groves of
Extremadura Extremadura (; ext, Estremaúra; pt, Estremadura; Fala: ''Extremaúra'') is an autonomous community of Spain. Its capital city is Mérida, and its largest city is Badajoz. Located in the central-western part of the Iberian Peninsula, it ...
, which are extensively cultivated for sporting purposes. The species introduced into the rivers of the Tagus basin are fish adapted to the middle and lower reaches of the Tagus river and its more abundant tributaries, reservoirs and ponds; their presence is mainly due to introductions for sporting purposes and is generally scattered.


Herpetos

Eighteen species of amphibians and four species of strictly aquatic reptiles have been recorded in the Tagus basin. Six species of urodele amphibians are distributed, the gallipato, ''Pleurodeles waltl'', is abundant in the basin, although it has populations threatened by isolation such as those in the south and northwest of the
Community of Madrid The Community of Madrid (; es, Comunidad de Madrid ) is one of the seventeen autonomous communities of Spain. It is located in the centre of the Iberian Peninsula, and of the Central Plateau (''Meseta Central''). Its capital and largest munici ...
. The fire salamander, ''Salamandra salamandra'', has populations threatened by isolation or habitat loss in Madrid (Miraflores de la Sierra, La Pedriza, Cercedilla, Pelayos de la Presa and Robledo de Chavela) and
Ávila Ávila (, , ) is a city of Spain located in the autonomous community of Castile and León. It is the capital and most populated municipality of the Province of Ávila. It lies on the right bank of the Adaja river. Located more than 1,130 m abov ...
(La Adrada and Sotillo de la Adrada). The alpine newt, ''Mesotriton alpestris'', introduced in the Peñalara Lagoon, the Iberian newt, ''Lissotriton boscai'', distributed throughout the basin except in the provinces of Province of Guadalajara, Guadalajara and Province of Cuenca, Cuenca, being very abundant in the
Sierra de Gredos The Sierra de Gredos is a mountain range in central Spain that spans the provinces of Ávila, Salamanca, Cáceres, Madrid, and Toledo. It is part of the much larger Sistema Central of mountain ranges. Its highest point is Pico Almanzor, at 2,5 ...
and the Tiétar valley. The marbled newt, ''Triturus marmoratus'', distributed in the
Central System The Central System, Spanish and pt, Sistema Central, is one of the main systems of mountain ranges in the Iberian Peninsula. The 2,592 m high Pico Almanzor is its highest summit. The Central System is located just north of the 40th parallel an ...
, the southern limit of its Iberian distribution; the Southern marbled newt, ''Triturus pygmaeus'', recently taxonomically separated from the marbled newt, continues the distribution of the latter towards the south, being, therefore, the one that occupies most of the basin. The number of species of anuran amphibians present in the basin is 12; the midwife toads present are the common, ''Alytes obstetricans'', located in the most northeastern area (Guadalajara and Cuenca mainly) and some areas of the Central System, southern limit of the distribution of the species) in the north of Cáceres, Ávila and Madrid. In these last locations it comes into contact with the Iberian midwife toad, ''Alytes cisternasii'', which is distributed towards the south, occupying a large part of the basin, where it is abundant. The Iberian painted frog, ''Discoglossus galganoi'', follow a pattern similar to that of the midwife toads. The Iberian spadefoot toad, ''Discoglossus galganoi'', is widely distributed in the western area; and the Spanish painted frog, ''Discoglossus jeanneae'', occupies the easternmost area of the basin (northwest of Madrid, Guadalajara and Cuenca, Spain, Cuenca) and its populations are threatened, depending on the state of conservation of the springs and fountains where they live. The Spanish spadefoot toad, ''Pelobates cultripes'', is distributed throughout the basin, being more abundant in the ''dehesas'' of the western area. It is a species considered to be in recession throughout the country and has numerous threatened populations, especially in the Community of Madrid. The Common parsley frog, ''Pelodytes punctatus'', is distributed in the basin mainly in its eastern zone, Madrid, Cuenca, Guadalajara and Toledo, and is considered a frequent but threatened species. The froglets of the Genus (biology), genus, ''Hyla'', the Mediterranean tree frog, ''Hyla meridionalis'' and the European tree frog, ''Hyla arborea'', are both distributed throughout the basin, with the European tree frog extending more towards the east and the Mediterranean tree frog being more abundant towards the southwest. They have an area of sympatry in the Tiétar river valley, where sterile Hybrid (biology), hybrid individuals have been located. The Iberian frog, ''Rana iberica'', occupies areas of the basin of a certain altitude, preferably above 2000 m a.s.l., in the
Central System The Central System, Spanish and pt, Sistema Central, is one of the main systems of mountain ranges in the Iberian Peninsula. The 2,592 m high Pico Almanzor is its highest summit. The Central System is located just north of the 40th parallel an ...
and in the Sierra de Guadalupe, Spain, Sierra de Guadalupe and Sierra de San Mamede in the south of Cáceres. The most abundant anurans in the territory of the Tagus basin are the common toad, ''Bufo bufo'' and the Natterjack toad, ''Bufo calamita'' and the Perez's frog, ''Rana perezi''. The group of reptiles adapted to aquatic environments present in the Tagus basin are the freshwater turtles with two species, the Spanish pond turtle, ''Mauremys leprosa'' and the European pond turtle, ''Emys orbicularis''. Of the European pond turtle there are populations in isolated nuclei and with small numbers of individuals in Madrid, in the basal floor of the
Sierra de Guadarrama The Sierra de Guadarrama (Guadarrama Mountains) is a mountain range forming the main eastern section of the Sistema Central, the system of mountain ranges along the centre of the Iberian Peninsula. It is located between the systems Sierra de G ...
and others even smaller in the western part of the
Central System The Central System, Spanish and pt, Sistema Central, is one of the main systems of mountain ranges in the Iberian Peninsula. The 2,592 m high Pico Almanzor is its highest summit. The Central System is located just north of the 40th parallel an ...
; on the southern slopes of the
Sierra de Gredos The Sierra de Gredos is a mountain range in central Spain that spans the provinces of Ávila, Salamanca, Cáceres, Madrid, and Toledo. It is part of the much larger Sistema Central of mountain ranges. Its highest point is Pico Almanzor, at 2,5 ...
, between Toledo and Province of Ávila, Avila and in some parts of the province of Cáceres. On the other hand, the best populations of Spanish pond turtle are found in Spain, being more abundant in the southern half of the country. This species is well distributed throughout the basin, being scarcer and more localized in the eastern area and abundant or very abundant in the middle and lower reaches of the main tributaries (Tiétar, Almonte, etc.) and other wetlands in the area. Another species of turtle existing in the basin is the introduced species, the Pond slider, ''Trachemys scripta''. The water snakes found in the basin are the viperine snake, ''Natrix maura'', and the ringed snake, ''Natrix natrix''. The viperine snake is a very common Snake, ophidian, using all types of aquatic environments in which it can have a high density of individuals. In contrast, the ringed snakes are more scarce and have fewer locations.


Birds

Within the group of birds, at least 53 breeding species have been cited in the area and which are closely linked to the riparian and/or helophytic vegetation of the rivers, lagoons and reservoirs of the basin. The following is a list of nesting birds and the birds included in the Annexes of the European Union, European Community Birds Directive, including two introduced
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
: During the winter, the number of birds that choose the aquatic biotypes of the Tagus to winter or that are passing through increases the ornithological population. In this way, birds such as the common crane, (''Grus grus''), the greylag goose, (''Anser anser''), ruff, (''Philomachus pugnax''), common snipe, (''Gallinago gallinago''), etc. can be observed. Also in the gorges, ravines and slopes of river terraces nest large numbers of cock-of-the-rock birds, some of them of great ecological value as the Eurasian griffon vulture (''Gyps fulvus''), Egyptian vulture (''Neophron percnopterus''), golden eagle (''Aquila chrysaetos''), Bonelli's eagle (''Hieraaetus fasciatus''), Eurasian eagle-owl (''Bubo bubo''), peregrine falcon (''Falco peregrinus''), etc.. Finally, the black stork (''Ciconia nigra''), which nests in the least disturbed and inaccessible areas of the Mediterranean forest, river cuts or mountain gorges, always in the vicinity of rivers, lagoons or reservoirs where it can feed.


Mammals

The mammal species directly related to the freshwater environment due to their aquatic or semi-aquatic nature in the Tagus basin are: the Eurasian otter (''Lutra lutra''), an autochthonous aquatic Mustelidae, mustelid adapted to this environment, distributed throughout the basin, except in unfavorable or polluted waters; another mustelid present is the American mink (''Mustela vison''), an allochthonous species, which has colonized some areas after escaping from farms or uncontrolled releases. The Pyrenean desman (''Galemys pyrenaicus'') is generally found above 700 m a.s.l., in streams and rivers of constant current and steep slope, in well oxygenated and clean waters. It is located in the headwaters of the Tagus tributaries, which originate in the Central System, in the provinces of Cáceres, Ávila, Madrid and Guadalajara. The Mediterranean water shrew (''Neomys anomalus''), an insectivorous mammal with excellent swimming and diving qualities, associated with permanent and well oxygenated watercourses and in areas of high humidity. Of the 23 species of bats that inhabit the Tagus basin, there is one that is closely linked to bodies of water, namely the Daubenton's bat (''Myotis daubentonii''), which feeds on invertebrates linked to the aquatic environment such as the winged phases of trichoptera, ephemeroptera, etc. Finally, the Southwestern water vole (''Arvicola sapidus''), which is linked to permanent aquatic environments and can be found up to 2100 m above sea level, is scattered throughout the basin.


Territorial distribution

The territorial scope of the Tagus basin, 55,645 km2 in the
Spanish Spanish might refer to: * Items from or related to Spain: **Spaniards are a nation and ethnic group indigenous to Spain **Spanish language, spoken in Spain and many Latin American countries **Spanish cuisine Other places * Spanish, Ontario, Cana ...
area, extends over five Autonomous communities of Spain, Autonomous Communities, totaling twelve provinces, as well as four Province, provincial capitals within the basin, Madrid, Toledo, Guadalajara, Spain, Guadalajara and Cáceres.


Hydraulic infrastructure

The National Water Works Plan of 1902, drawn up by the Corps of Engineers, included the works necessary for the irrigation of 181,850 ha in the Tagus basin. In 1933, the year in which a new Plan was drawn up, practically none of the previously programmed works had been carried out, only the reservoirs of El Burguillo Reservoir, El Burguillo and El Charco del Cura Reservoir, El Charco del Cura, with an eminently hydroelectric purpose, achieved a regulation of the
Alberche The Alberche is a river in the provinces of Ávila, Madrid and Toledo, central Spain. It begins its course at 1,800 m in Fuente Alberche, San Martín de la Vega del Alberche municipal term, Ávila Province. It forms the natural division between ...
river and could allow irrigation in the area of Talavera de la Reina. This new Plan included the works not previously carried out, reducing the irrigable surface area to 110,000 ha, a limitation imposed by the projected transfer to the Levante area of the resources which, according to the Plan, were supposed to be surplus in the basin; from then on, the planning of the possible uses began, which, interrupted by the Spanish Civil War of 1936, began to be developed from 1940 onwards.


Tagus-Segura Diversion

The first time that the Tagus basin was considered as part of a solution to the problem of scarcity of water resources in the southeast was in Spain in the 1930s. From 1960 onwards, studies of water resources were systematically carried out and the potential for irrigation and supply were evaluated. The transfer infrastructures are, essentially, the following: * Two regulating reservoirs, Entrepeñas Reservoir, Entrepeñas and Buendía Reservoir, Buendía, with a capacity of 835 and 1639 hm³, connected by a tunnel that allows flows from the former to the latter to be evacuated. * The .Bolarque Reservoir, Bolarque reservoir, with a capacity of 31 hm³, where the intake is located and from where the water is distributed. * The elevation from Bolarque to the small reservoir of La Bujeda Reservoir, La Bujeda, sized to provide, in coordination with Bolarque, the hydroelectric regulation. It consists of four groups capable of lifting 66 m³ at a height of 243 m, with a maximum power of 208 Watt#Multiples, MW. The plant is reversible, the normal operation consists of lifting to La Bujeda and turbining from La Bujeda to Bolarque to produce electricity, 400 MegaWatt, MW. * The canal that starts at La Bujeda is capable of transporting 33 m³/s and consists of two parts; from La Bujeda to the tail of the Alarcón Reservoir, Alarcón reservoir, in the Júcar basin, and another from the outlet of the latter to the Talave reservoir, in the Segura basin, with respective lengths of 93 and 135 km for each section; Along this route, there are several Aqueduct (water supply), aqueducts, the most outstanding being the Cigüela aqueduct, which is 6190 m long and 44 m high, several tunnels, 12 in the first section with a total of 11,878 m, and the Talave tunnel in the second section, 32 km long. There are also other singular works, such as the Altomira balancing chimney, 69 m high, and several Hydroelectricity, hydroelectric power production plants.


Events

Table with episodes of meteorological drought that occurred in the
Tagus The Tagus ( ; es, Tajo ; pt, Tejo ; see #Name, below) is the longest river in the Iberian Peninsula. The river rises in the Montes Universales near Teruel, in mid-eastern Spain, flows , generally west with two main south-westward sections ...
demarcation up to the end of the 19th century.


Large floods

The rivers of the Tagus demarcation have generated, prior to their regulation, floods that have often resulted in major floods, causing great damage, causing personal and material losses. The areas with the greatest number of floods were: the areas of the Tagus river in Talavera de la Reina and
Aranjuez Aranjuez () is a city and municipality of Spain, part of the Community of Madrid. Located in the southern end of the region, the main urban nucleus lies on the left bank of Tagus, a bit upstream the discharge of the Jarama. , the municipality h ...
, the
Jarama Jarama () is a river in central Spain. It flows north to south, and passes east of Madrid where the El Atazar Dam is built on a tributary, the Lozoya River. It flows into the river Tagus in Aranjuez. The Manzanares is a tributary of the Jarama ...
river in San Fernando de Henares, the Tiétar river and the Alagón river.


Portuguese Tagus

The surface area of the Portuguese part of the basin accounts for more than 28% of the continental part of Portugal, including entirely the districts of Santarém District, Santarém and Castelo Branco, Portugal, Castelo Branco and a significant part of those of Lisbon District, Lisbon, Leiria District, Leiria, Portalegre District, Portalegre, Guarda District, Guarda, Évora District, Évora and Setúbal District, Setúbal. It includes all or part of 94 "''Concelhos''", with an area of more than 30,000 km2, where about 3.5 million inhabitants live (more than one third of the country's population). It is made up of 16 hydrographic sub-basins corresponding to the main tributaries of the Tagus River, a small endorheic basin and the small basins that drain into the Paja Sea and the Atlantic Ocean, between the Costa da Caparica and Cabo Espichel.


Alenquer sub-basin

Small sub-basin on the right bank of the Tagus, with 282 km2, partially comprising the municipalities of Alenquer, Portugal, Alenquer, Azambuja and Cadaval. Composed of two distinct Lithology, lithological groups, to the west calcareous rocks, fractured and karsified landscapes, with essentially subway runoff and the other group represented by recent rocks,
Cenozoic The Cenozoic ( ; ) is Earth's current geological era, representing the last 66million years of Earth's history. It is characterised by the dominance of mammals, birds and flowering plants, a cooling and drying climate, and the current configura ...
, sandy, very permeable in its great majority. The flora species, endemic, rare or vulnerable, linked to the edaphic-hygrophilous environments: ''Glinus lotoides, Euphorbia uliginosa, Myosotis lusitanica'' and ''Juncus valvatus''. The valleys have inaccessible margins covered with well-preserved shrub and tree vegetation. The presence of Iberian emerald lizards in some of the riverbanks associated with agricultural valleys, with their fragmented distribution and the degradation of the riverbanks, are two aspects to consider for the conservation of the species.


Almonda sub-basin

Small sub-basin on the right bank of the Tagus, 274 km2, including the municipalities of Alcanena, Entroncamento, Golegã, Golega, Porto de Mós, Santarém, Portugal, Santarém and Torres Novas. It consists of fractured calcareous rocks, with eminently subway Surface runoff, runoff, contributing as a recharge structure for the free, semi-confined and confined aquifers in the area. Within the zone there are several areas of special protection: * Serras de Aire e Candeeiros Natural Park. * Boquilobo Bog Natural Reserve. * Paul do Boquilobo Special Protection Zone. The most important area of the sub-basin is Paul do Boquilobo, whose flora has been very degraded in recent years, and some of the species identified there 20 years ago have not been observed recently. The species that settle there are ''Oenanthe fistulosa, Butomus umbellatus, Damasonium alisma'' and ''Galium palustre.''


Alviela sub-basin

Sub-basin of 331 km2, on the right bank of the Tagus, in the municipalities of Idanha-a-Nova and Castelo Branco, Portugal, Castelo Branco. Sub-basin with sedimentary cover, with a granitic outcrop located on its eastern edge. The flora species present are ''Salix salviifolia, Salix salvifolia'' subsp. ''australis'' and the importance of the rocky escarpments for rupicolous birds, especially for nocturnal and diurnal birds of prey and the black stork. In these areas mammals can be found that use the watercourses and their margins as a means of movement between territories (''European wildcat, Iberian wolf, Iberian lynx, otter,'' etc.).


Erges sub-basin

The sub-basin of the Erges (river), Erges river, which forms a border with Spain in a large part of its course, with an area of 593 km2 in the municipalities of Idanha-a-Nova and Penamacor. Lithologically dominated by impermeable rocks, quartzite ridges and a recent sedimentary cover (Cenozoic), which can acquire characteristics of a free aquifer. In this sub-basin, the rocky escarpments play an important role, where a large number of rupicolous birds nest, and it is included in the Special Protection Zone of the International Tagus.


Zézere sub-basin

The Zêzere River, Zézere river basin is one of the largest sub-basins of the Tagus river in Portugal, with 5076 km2 (about 20% of the total). It includes the municipalities of Alvaiázere, Belmonte, Portugal, Belmonte, Castanheira de Pera, Ferreira do Zêzere, Sertã and Vila de Rei and part of twenty-four others. It extends over an extensive area of the northernmost part of the basin, dominated by granitic outcrops in the southern part of this sub-basin, originating narrow valleys. The flora, endemic or vulnerable linked to hygrophilous environments are: ''Osmunda regalis, Montia fontana subsp. amporitana, Glinus lotoides, Gratiola linifolia, Ludwigia palustris''. This area is rich and varied from the point of view of habitat that enhances the quality of the fauna it shelters, so in the upper courses of the Zézere and its tributaries, the species of ''Chioglossa lusitanica'' and ''Galemys pyrenaicus'' stand out. In the middle part of the fluvial course, they stand out for the nesting of rupicolous birds such as the ''Eurasian eagle-owl''.


Notes


References

{{reflist


External links


Tagus Hydrographic Confederation
(in Spanish)

(in Spanish)
Spanish Society of Dams and Reservoirs
(in Spanish)
Geographic viewer of the SIA. Map of large river basins
(in Spanish)
Digital Water Book. Surface Water Bodies. Rivers
(in Spanish) Tagus Tagus basin Basins Drainage basins Geology of Spain Basins of Europe